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All are animals divided into segments. Centipedes, most of which are carnivores, have a pair of legs on each segment, and millipedes have two pairs of legs per segment. These invertebrates (which are not insects) have so many legs that, to walk, they must use a highly sophisticated timing mechanism that seems to follow mathematical principles. Applied Math A Thousand Legs To walk, land-dwelling arthropods arch their bodies and move their six legs in coordination so that when one leg moves forward in a power stroke, the ones in front, behind, and opposite it are in a recovery phase, remaining on the ground.

One of its characteristics is the nymph stage of young insects. The nymph gradually changes in shape as it grows. When it sheds its exoskeleton, the adult emerges. 3 IMAGO (ADULT) INSIDE THE LARVA EMPEROR DRAGONFLY Anax imperator 1 EGG The insect's heart, nervous system, and breathing system are almost completely developed during the larval stage, and they change very little afterward. The reproductive system is formed later. 2 NYMPH FAREWELL TO THE OLD BODY INTESTINE The larva's last exoskeleton begins to fall off and is replaced by a greenish tissue that will form the cocoon, or chrysalis.

5 million, are a testimony to their evolutionary success. They have been successful, in part, because they are small, need less food than larger organisms, and have extraordinarily developed means of movement that keep them from being easy victims for predators. 54 INSECTS INVERTEBRATES 55 The Secret of Success Taking Many Steps The class Chilopoda (centipedes, including many carnivores and predators) and the class Diplopoda (millipedes) are known as myriapods. Their patterns of movement are both complex and efficient.

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